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Recently [ when? Whereas in the mids, only Pseudopodoces , Baeolophus , Melanochlora , and Sylviparus were considered well-supported by the available data as distinct from Parus.

In the Sibley-Ahlquist taxonomy , the family Paridae is much enlarged to include related groups such as the penduline tits and long-tailed tits , but while the former are quite close to the tits and could conceivably be included in that family together with the stenostirid "warblers" , the long-tailed tits are not.

Indeed, the yellow-browed tit and the sultan tit are possibly more distant to the tits than the penduline tits are.

Alternatively, all tits—save the two monotypic genera discussed in the preceding section and possibly Cyanistes , but including Hume's ground tit—could be lumped in Parus.

In any case, four major clades of "typical" tits can be recognized: the dark-capped chickadees and their relatives Poecile including Sittiparus , the long-crested Baeolophus and Lophophanes species, the usually tufted, white-cheeked Periparus including Pardaliparus with more subdued coloration and finally Parus sensu stricto including Melaniparus and Machlolophus.

Still, the interrelationship of these, as well as the relationships of many species within the clades, are not well-resolved at all; analysis of morphology and biogeography probably gives a more robust picture than the available molecular data.

The first were the ancestors of Baeolophus , with chickadees arriving somewhat later. Cephalopyrus flammiceps. Sylviparus modestus. Melanochlora sultanea.

Parus monticolus. From Wikipedia, the free encyclopedia. Family of small passerine birds. For other uses, see Tit. Online Etymology Dictionary.

Retrieved Forshaw, Joseph ed. Encyclopaedia of Animals: Birds. London: Merehurst Press. Handbook of the Birds of the World.

Volume Picathartes to Tits and Chickadees. Barcelona: Lynx Edicions. Integrative and Comparative Biology. M; Farine, D. Ecology Letters.

Retrieved 7 December Proc Biol Sci. Christopher Helm , London. Species relationships based on sequences of the mitochondrial cytochrome-b gene".

P Molecular Phylogenetics and Evolution. World Bird List Version 6. International Ornithologists' Union. Retrieved 15 February Pseudopodoces humilis , a misclassified terrestrial tit Paridae of the Tibetan Plateau: evolutionary consequences of shifting adaptive zones.

Ibis — Hidden categories: All articles with dead external links Articles with dead external links from February Webarchive template wayback links Articles with short description Short description is different from Wikidata Articles with 'species' microformats All articles with vague or ambiguous time Vague or ambiguous time from May Commons category link is on Wikidata.

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Wikimedia Commons Wikispecies. European crested tit in Scotland. Paridae Vigors , The great tit is a distinctive bird with a black head and neck, prominent white cheeks, olive upperparts and yellow underparts, with some variation amongst the numerous subspecies.

It is predominantly insectivorous in the summer, but will consume a wider range of food items in the winter months, including small hibernating bats.

The female lays around 12 eggs and incubates them alone, although both parents raise the chicks. In most years the pair will raise two broods.

The nests may be raided by woodpeckers , squirrels and weasels and infested with fleas , and adults may be hunted by sparrowhawks.

The great tit has adapted well to human changes in the environment and is a common and familiar bird in urban parks and gardens.

The great tit is also an important study species in ornithology. The great tit was described under its current binomial name by Carl Linnaeus in his 10th edition of Systema Naturae.

The great tit was formerly treated as ranging from Britain to Japan and south to the islands of Indonesia, with 36 described subspecies ascribed to four main species groups.

The major group had 13 subspecies across Europe, temperate Asia and north Africa, the minor group's nine subspecies occurred from southeast Russia and Japan into northern southeast Asia and the 11 subspecies in the cinereus group were found from Iran across south Asia to Indonesia.

The three bokharensis subspecies were often treated as a separate species, Parus bokharensis , the Turkestan tit. This form was once thought to form a ring species around the Tibetan Plateau , with gene flow throughout the subspecies, but this theory was abandoned when sequences of mitochondrial DNA were examined, finding that the four groups were distinct monophyletic and that the hybridisation zones between the groups were the result of secondary contact after a temporary period of isolation.

A study published in confirmed that the major group was distinct from the cinereus and minor groups and that along with P.

The divergence between the bokharensis and major groups was estimated to have been about half a million years ago.

The study also examined hybrids between representatives of the major and minor groups in the Amur Valley where the two meet.

Hybrids were rare, suggesting that there were some reproductive barriers between the two groups. The study recommended that the two eastern groups be split out as new species, the cinereous tit Parus cinereus , and the Japanese tit Parus minor , but that the Turkestan tit be lumped in with the great tit.

The nominate subspecies of the great tit is the most widespread, its range stretching from the Iberian Peninsula to the Amur Valley and from Scandinavia to the Middle East.

The other subspecies have much more restricted distributions, four being restricted to islands and the remainder of the P. The dominance of a single, morphologically uniform subspecies over such a large area suggests that the nominate race rapidly recolonised a large area after the last glacial epoch.

This hypothesis is supported by genetic studies which suggest a geologically recent genetic bottleneck followed by a rapid population expansion.

The genus Parus once held most of the species of tit in the family Paridae , but morphological and genetic studies led to the splitting of that large genus in The great tit was retained in Parus , which along with Cyanistes comprises a lineage of tits known as the "non-hoarders", with reference to the hoarding behaviour of members of the other clade.

The genus Parus is still the largest in the family, but may be split again. Hybrids with tits outside the genus Parus are very rare, but have been recorded with blue tit , coal tit , and probably marsh tit.

There are currently 15 recognised subspecies of great tit: [10]. The great tit is large for a tit at The nominate race P. The breast is bright lemon-yellow and there is a broad black mid-line stripe running from the bib to vent.

There is a dull white spot on the neck turning to greenish yellow on the upper nape. The rest of the nape and back are green tinged with olive.

The wing-coverts are green, the rest of the wing is bluish-grey with a white wing-bar. The tail is bluish grey with white outer tips.

The plumage of the female is similar to that of the male except that the colours are overall duller; the bib is less intensely black, [10] as is the line running down the belly, which is also narrower and sometimes broken.

There is some variation in the subspecies. The plumage of P. It is also slightly smaller, with a smaller bill but longer tail.

The situation is similar for the two related subspecies in the Turkestan tit group. The colour of the male bird's breast has been shown to correlate with stronger sperm, and is one way that the male demonstrates his reproductive superiority to females.

Higher levels of carotenoid increase the intensity of the yellow of the breast its colour, and also enable the sperm to better withstand the onslaught of free radicals.

The great tit is, like other tits, a vocal bird , and has up to 40 types of calls and songs. The calls are generally the same between the sexes, but the male is much more vocal and the female rarely calls.

Soft single notes such as "pit", "spick", or "chit" are used as contact calls. A loud "tink" is used by adult males as an alarm or in territorial disputes.

One of the most familiar is a "teacher, teacher", often likened to a squeaky wheelbarrow wheel, which is used in proclaiming ownership of a territory.

One explanation for the great tit's wide repertoire is the Beau Geste hypothesis. The eponymous hero of the novel propped dead soldiers against the battlements to give the impression that his fort was better defended than was really the case.

Similarly, the multiplicity of calls gives the impression that the tit's territory is more densely occupied than it actually is. Whether the theory is correct or not, those birds with large vocabularies are socially dominant and breed more successfully.

The great tit has a wide distribution across much of Eurasia. It can be found across all of Europe except for Iceland and northern Scandinavia , including numerous Mediterranean islands.

The great tit occupies a range of habitats. It is most commonly found in open deciduous woodland , mixed forests , forest edges and gardens.

In dense forests, including conifer forests it prefers forest clearings. In northern Siberia it lives in boreal taiga.

In North Africa it rather resides in oak forests as well as stands of Atlas cedar and even palm groves. In the east of its range in Siberia, Mongolia and China it favours riverine willow and birch forest.

Riverine woodlands of willows, poplars are among the habitats of the Turkestan subspecies, as well as low scrubland , oases ; at higher altitudes it occupies habitats ranging from dense deciduous and coniferous forests to open areas with scattered trees.

The great tit is generally not migratory. Pairs will usually remain near or in their territory year round, even in the northern parts of their range.

Young birds will disperse from their parents' territory, but usually not far. Populations may become irruptive in poor or harsh winters, meaning that groups of up to a thousand birds may unpredictably move from northern Europe to the Baltic and also to Netherlands, Britain, even as far as the southern Balkans.

The great tit was unsuccessfully introduced into the United States; birds were set free near Cincinnati, Ohio between and but failed to become established.

Suggestions that they were an excellent control measure for codling moths nearly led to their introduction to some new areas particularly in the United States of America, however this plan was not implemented.

Great tits are primarily insectivorous in the summer, feeding on insects and spiders which they capture by foliage gleaning. Seeds and fruit usually come from deciduous trees and shrubs, like for instance the seeds of beech and hazel.

Where it is available they will readily take table scraps, peanuts and sunflower seeds from bird tables. Large food items, such as large seeds or prey, are dealt with by "hold-hammering", where the item is held with one or both feet and then struck with the bill until it is ready to eat.

Using this method, a great tit can get into a hazelnut in about twenty minutes. When feeding young, adults will hammer off the heads off large insects to make them easier to consume, and remove the gut from caterpillars so that the tannins in the gut will not retard the chick's growth.

Great tits combine dietary versatility with a considerable amount of intelligence and the ability to solve problems with insight learning, that is to solve a problem through insight rather than trial and error.

This is the first time a songbird has been recorded preying on bats. The tits only do this during winter when the bats are hibernating and other food is scarce.

Great tits are monogamous breeders and establish breeding territories. Females are likely to disperse to new territories if their nest is predated the previous year.

If the pair divorces for some reason then the birds will disperse, with females travelling further than males to establish new territories.

Great tits are seasonal breeders. The exact timing of breeding varies by a number of factors, most importantly location.

Most breeding occurs between January and September; in Europe the breeding season usually begins after March. In Israel there are exceptional records of breeding during the months of October to December.

The amount of sunlight and daytime temperatures will also affect breeding timing. Great tits are cavity nesters, breeding in a hole that is usually inside a tree, although occasionally in a wall or rock face, and they will readily take to nest boxes.

The nest inside the cavity is built by the female, and is made of plant fibres, grasses, moss, hair, wool and feathers. The number in the clutch is often very large, as many as 18, but five to twelve is more common.

Clutch size is smaller when birds start laying later, and is also lower when the density of competitors is higher. Insularity also affects clutch size, with great tits on offshore islands laying smaller clutches with larger eggs than mainland birds.

The female undertakes all incubation duties, and is fed by the male during incubation. The timing of hatching, which is best synchronised with peak availability of prey, can be manipulated when environmental conditions change after the laying of the first egg by delaying the beginning of incubation, laying more eggs or pausing during incubation.

The chicks, like those of all tits , are hatched unfeathered and blind. Once feathers begin to erupt, the nestlings are unusual for altricial birds in having plumage coloured with carotenoids similar to their parents in most species it is dun-coloured to avoid predation.

The nape is yellow and attracts the attention of the parents by its ultraviolet reflectance. This may be to make them easier to find in low light, or be a signal of fitness to win the parents' attention.

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